Lecithoceridae (Lepidoptera) of Madagascar: I. Descriptions of eighteen new species of Torodora Meyrick, 1894.

The present study is aimed at describing lecithocerid specimens from Madagascar collected by Pierre Viette and his colleagues during the 1950's and preserved in the Muséum National d'Histoire Naturelle (MNHN, Paris). Eighteen new species of Torodora Meyrick, 1894 are described, namely: T. hana Park & Koo, sp. nov., T. larseni Park & Koo, sp. nov., T. toliarensis Park & Koo, sp. nov., T. silvestris Park & Koo, sp. nov., T. sogai Park & Koo, sp. nov., T. vietteiola Park & Koo, sp. nov., T. haploista Park & Koo, sp. nov., T. septentriella Park & Koo, sp. nov., T. zahamenensis Park & Koo, sp. nov., T. unicolorella Park & Koo, sp. nov., T. pistillia Park & Minet, sp. nov., T. andringitrensis Park & Minet, sp. nov., T. ochrizona Park & Minet, sp. nov., T. anosibensis Park & Minet, sp. nov., T. convexula Park & Minet, sp. nov., T. sphenosa Park & Minet, sp. nov., T. malagasiella Park & Minet, sp. nov., and T griveaudi Park & Minet, sp. nov. The previously unknown male of T. lecithocerella (Viette, 1956) is described for the first time. Adults and their male or female genitalia are illustrated.

Review of the genera Thubana Walker and Torodora Meyrick (Lepidoptera: Lecithoceridae) from Sumatra, Indonesia.

Sumatra is the second largest island that is fully within Indonesian territory, but it is one of the most poorly explored areas for species diversity of the family Lecithoceridae, with less than 10 known species. In the present paper, six new species of Thubana Walker, 1864 (T. sumatrana sp. nov., T. spiniosa sp. nov., T. lata sp. nov., T. prapatensis sp. nov., T. siantarensis sp. nov., and T. selenisa sp. nov.) and three new species of Torodora Meyrick (T. diehliella sp. nov., T. exilivalvata sp. nov., and T. squariella sp. nov.) are described from Sumatra. In addition, Thubana ochracea Park & Abang, 2005, which was described from Sarawak, Malaysia, is reported for the first time from Sumatra. Illustrations of adults and genitalia of all new species are given. A list of the known species of the subfamily Torodorinae from Indonesia is provided.

Review of the genus Lecithocera Herrich-Schffer (Lepidoptera, Lecithoceridae) in DR Congo, with description of four new species.

The genus Lecithocera Herrich-Schffer, 1853 in the Democratic Republic of theCongo (DR Congo) is reviewed, with descriptions of four new species: Lecithocera katangaica Park, sp. nov., L. seydeliella Park, sp. nov., L. cornutispina Park, sp. nov., and L. lusingensis Park, sp. nov. Five species are already recorded from this country: 1) L. chloroscia Meyrick described from DR Congo; 2) L. lucernata Meyrick previously reported from DR Congo by Ghesquire in 1940; 3) L. trifera Meyrick and 4) L. xanthochalca Meyrick, their generic assignments still remain doubtable; and 5) L. ideologa Meyrick. Lecithocera dicentropa Meyrick, 1938 is synonymized as a junior synonym of L. ideologa Meyrick. Adults and the genitalia of the new species and L. ideologa Meyrick are illustrated, and a lectotype for L. trifera Meyrick is designated herein.

The family Lecithoceridae (Lepidoptera: Gelechioidea) from Ivory Coast (Republic of Cte dIvoire), with descriptions of nine new species.

The family Lecithoceridae is reviewed with descriptions of nine new species from Ivory Coast (Republic of Cte dIvoire): four new species of Thubdora Park, 2018 (T. ivoryensis Park Karisch, sp. nov., T. pulchella Park Karisch, sp. nov., T. tonkpiensis Park Karisch, sp. nov., and T. cuneiformis Park Karisch, sp. nov.); and four new species of Dragmatucha Meyrick, 1908 (D. polystriana Park Karisch, sp. nov., D. mediolinea Park Karisch, sp. nov., D. ovartiella Park Karisch, sp. nov., and D. flavoaurea Park Karisch, sp. nov.) of the subfamily Torodorinae, and one new species of Pectinimura Park, 2008 (P. paraliberalis Park Karisch, sp. nov.) of the subfamily Lecithocerinae. In addition, Thubdora barbata (Meyrick, 1933) and Thubdora vernaculella Park Karisch, 2021 are reported for the first time from Ivory Coast, and the female of T. barbata is described for the first time. Images for adults and the genitalia of all species are provided.

Description of Spiniola hanaro Park, gen. and sp. nov., and three new species of Torodora Meyrick, 1894 (Lepidoptera, Lecithoceridae, Torodorinae) from the Afrotropical Region.

A new genus, Spiniola Park, gen. nov., is described from Malawi, based on the type species, S. hanaro Park, sp. nov., and three new species of Torodora Meyrick, 1894 are described: T. batillana Park Koo, sp. nov. from Kenya, and T. chrysotes Park Koo, sp. nov. and T. manalis Park Koo, sp. nov. from Uganda. Images of adults and genitalia of the new species are provided.

Two new species of Gelechiidae (Lepidoptera) from Korea, with some biological data including larval host plants.

In this study, two new species of the family Gelechiidae are described from Korea. The new species are Teleiodes juglansivora Park Byun, sp. nov. and Encoplata najuensis Park Byun, sp. nov. For the new species, some biological data concerning on their larval food plants are given.

The family Lecithoceridae (Lepidoptera, Gelechioidea) from Bioko Island (Equatorial Guinea), with descriptions of five new species.

Five new species belonging to the subfamily Torodorinae of Lecithoceridae are described from Bioko Island (Equatorial Guinea), based on specimens which were collected during a short expedition on the island by the second author in 1994. The new species are three species of the genus Thubdora Park, 2018 (T. biocoica sp. nov., T. vernaculella sp. nov., and T. angustiala sp. nov.), and a new species of Ptilothyris Walsingham, 1891and Dragmatucha Meyrick, 1908 respectively (Ptylothyris ruicheensis sp. nov.; and Dragmatucha vulcaniella sp. nov.). In addition, Thubdora bilobella Park, 2018 is reported for the first time from Equatorial Guinea. Images for adults and the genitalia of all species are given.

Four new species of Thubdora Park, 2018 and one new species of Torodora Meyrick, 1894 (Lepidoptera, Lecithoceridae, Torodorinae) from Tanzania.

Four new species of Thubdora Park, 2018 and one new species of Torodora Meyrick,1894 are described from Tanzania. The new species are Thubdora forficatalis Park Cho, sp. nov., T. mufindiensis Park Cho, sp. nov., T. ochrospilosa Park Cho, sp. nov., T. tanzaniana Park Cho, sp. nov., and Torodora melanonota Park Cho, sp. nov. Images of adults and genitalia of the new species are provided and a map for the type localities of the new species is given.

A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species.

A new genus Viperinus Park, gen. nov., belonging to Torodorinae (Lecithoceridae), is proposed, with descriptions of two new species, V. orbiosus Park Koo sp. nov. from Kenya and V. pyknoistus Park Koo, sp. nov. from Uganda. Furthermore, two new species of Protolychnis Meyrick, 1925, P. oculiella Park Koo, sp. nov. and P. morogorensis Park Koo, sp. nov., are described from Kenya and Tanzania, respectively. In addition, P. petiliella Park, 2020 is reported for the first time from Tanzania. The new genus, Viperinus is very similar to Protolychnis in the superficial characters, but it is separated from the latter by having unique characters in genitalia: a pair of processes arising from base of each valva in the male genitalia and a polygonal signal plate in the female genitalia. Images of adults and genitalia of the four new species are provided, and a checklist of Protolychnis Meyrick is also given.

A new species of Tisis Walker from North Sumatra, Indonesia (Lepidoptera: Lecithoceridae).

A new species of the genus Tisis Walker is described from Sumatra, Indonesia, which is very close to T. sumatraensis Park, 2007.

Two new species records of Lecithoceridae (Gelechioidea: Lepidoptera) from India.

The Dalma Wildlife Sanctuary, located in the Dalma Hills about 10 km north of the city of Jamshedpur, Jharkhand, India, is topographically and climatically unique. It encompasses a forested area of over 195 km2 east of the Singhbhum and Saraikela-kharsawan districts of Jharkhand, extending from Chandil about 40 km eastward, running nearly parallel to the National Highway-33 at an altitude of about 280 m. The forests of the sanctuary are "dry peninsular sal" and "northern dry mixed deciduous forest".

Two new species of the genus Thubdora Park and a new species of Ptilothyris Walsingham (Lepidoptera, Gelechioidea, Lecithoceridae) from Uganda.

Two species of Thubdora Park (T. onsemiro Park, sp. nov. and T. elgozmanyi Park, sp. nov.) and one species of Ptilothyris Walsingham (P. galbiplatella Park, sp. nov.) are described from Uganda.

Review of the Oriental genus Pompographa Gozmány (Lepidoptera, Lecithoceridae), with description of a new species from Thailand.

A small Oriental genus Pompographa Gozmány, 1971 is reviewed, with three known species including a new species P. concaviella Park, sp. nov. described from Thailand. The recently described species from Laos, Torodora barathrona Park, 2020 is transferred to Prompographa, P. barathrona Park, comb. nov. Adult images of the three known species and genitalia of the two species, except the type species, are given.

Five new species of the genus Torodora Meyrick (Lepidoptera, Lecithoceridae, Torodorinae) from Cambodia, with a tentative check-list of the genus.

Five new species of the genus Torodora Meyrick of the family Lecithoceridae are described from Cambodia. The new species are T. helvinotula Park, sp. nov., T. spathiana Park, sp. nov., T. alterniella Park, sp. nov., T. triquetrella Park, sp. nov., and T. cupriella Park, sp. nov.. In addition, Torodora macrosigna Gozmány, 1973 is reported for the first time from Cambodia, with illustration of the male genitalia for the first time.

Two new genera, Halista Park, gen. nov. and Mireana Park, gen. nov. and six new species of the family Lecithoceridae (Lepidoptera) from the Philippines.

Two new genera, Halista Park, gen. nov. (type species: Ephalista batillosa Park Kim, sp. nov.) and Mireana Park, gen. nov. (type species: Miraena tawitawiensis Park Kim sp. nov.), and four new species, assigned to the previously known genera: Lecithocera wolframmeyi sp. nov, L. bipunctata sp. nov., Torodora platiella sp. nov. and Chrysonasma occidentis sp. nov. are described from the Philippines. Illustrations of adults and genitalia of all new species are provided.

Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam.

Twelve new species of the genus Dichomeris Hübner, 1818 and one new species of the genus Helcystogramma Zeller, 1877 of the subfamily Dichomeridinae of Gelechiidae are described from Vietnam, in combination with Chinese specimens. The new species are D. fascialobella Park Li, sp. nov., D. flavimaculata Park Li, sp. nov., D. infundibularis Park Li, sp. nov., D. lancea Park Li, sp. nov., D. longivalvata Park Li, sp. nov., D. marginalis Park Li, sp. nov., D. myriaodontosa Park Li, sp. nov., D. ochriella Park Li, sp. nov., D. quadratella Park Li, sp. nov., D. sclerospinula Park Li, sp. nov., D. spinosella Park Li, sp. nov., D. unilobata Park Li, sp. nov., and Helcystogramma albimacularis Park Li, sp. nov. In addition, eighteen Dichomeris species and three Helcystogramma species are reported for the first time from Vietnam, and of these, four previously unknown female are firstly found and described: Dichomeris parvisexafurca Li, Zhen Kendrick, 2010, Dichomeris argentenigera Li, Zhen Kendrick, 2010, Dichomeris splendiptera Li Park, 2017 and Dichomeris parallelosa Park Ponomarenko, 1998. Images of adults and genitalia for the new species and newly reported species including the firstly described females are provided.

A taxonomic review of the Afrotropical genus Dragmatucha Meyrick, 1908 (Lepidoptera, Gelechioidea, Lecithoceridae, Torodorinae), with descriptions of eleven new species.

The genus Dragmatucha Meyrick is reviewed and 11 species of a total of 13 are described as new. The new species originate from the Democratic Republic of Congo, Ghana, Kenya, and Uganda. Based on morphological characters, especially the color pattern of both wings, the genus is divided into two species-groups: the proaula species-group with five species (D. proaula Meyrick, 1908; D. galbinea Park, sp. nov.; D. kabarolensis Park, sp. nov.; D. saltualis Park, sp. nov. and D. vittatella Park, sp. nov.) and the hispidula species-group with eight species (D. hispidula Park, 2018; D. goniotes Park, sp. nov.; D. cochliana Park, sp. nov.; D. pedalis Park, sp. nov.; D. ghanaensis Park, sp. nov.; D. crinifrutalis Park, sp. nov.; D. dizostera Park, sp. nov. and D. kakumensis Park, sp. nov.). Adults and male genitalia of all new species are illustrated, but female genitalia are provided only for D. ghanaensis Park, sp. nov. and D. kakumensis Park, sp. nov. When fresh material was available, the analysis of COI sequences was undertaken.

Genus Torodora Meyrick (Lepidoptera, Lecithoceridae, Torodorinae) in Laos, with descriptions of five new species and two newly recorded species.

Genus Torodora Meyrick of the family Lecithoceridae in Laos is reviewed, describing five new species and reporting two unrecorded species of the genus. The new species are T. barathrona Park, sp. nov., T. canalis Park, sp. nov., T. cordisigna Park, sp. nov., T. digitalis Park, sp. nov., and T. multispinata sp. nov. In addition, T. metasaris (Meyrick, 1911), comb. nov. and Torodora pegasana Wu Liu, 1994 are reported for the first time from Laos. Images of adults and genitalia for the new species and the first known species are provided, along with a list of the previously known species with new collected localities.

Review of the Malagasy lecithocerid species described by Pierre Viette and deposited in MNHN (Paris), with new generic combinations and descriptions of a new subfamily and genus of Momphidae (Lepidoptera: Gelechioidea).

The holotypes of the Malagasy Lecithoceridae described by Pierre Viette (from 1954 to 1988) are reviewed, which represents a total of 23 specimens originally assigned to the genera Idiopteryx Walsingham, 1891 (4 named species) and Lecithocera Herrich-Shäffer, 1853 (19 named species). The holotype of Nemophora janineae Viette, 1954, erroneously described in Adelidae, is also included in the present study since it clearly belongs to the Lecithoceridae; N. janineae is here transferred to the genus Parkiana Cho, 2020. All these type specimens are housed in the Muséum National d'Histoire Naturelle (MNHN, Paris), where Pierre Viette had worked as a researcher during almost forty years (1945-1982). The Lecithocera species in question are transferred either to other genera of the Lecithoceridae, or to other families: 12 species, including Lecithocera kambanella and L. masoalella previously transferred by Park (2018), are assigned to Torodora Meyrick, 1894; three species to Parkiana Cho; the remaining species to non-lecithocerid groups, namely the Gelechiidae: Dichomeridinae (1 species), the Epichostis-group (considered incertae sedis but here expanded to include the genus Merocrates Meyrick, 1931; 1 species), and the Momphidae (2 described species, here regarded as synonyms, viz. L. andrianella Viette, 1968 and L. ranavaloella Viette, 1968). The four alleged Malagasy Idiopteryx turn out to belong to non-lecithocerid families and are transferred to the genera Dichomeris Hübner (Gelechiidae: Dichomeridinae; 1 species), Odites Walsingham (Peleopodidae: Oditinae; 1 species), and Moca Walker (Immidae, superfamily Immoidea; 2 species). Within the Momphidae, Lecithocera andrianella Viette is assigned to a new genus, Adelomompha Minet Park gen. nov., which is placed in a new subfamily (Adelomomphinae Minet Park subfam. nov.) since it lacks at least two imaginal apomorphies present in the ground plan of the remaining Momphidae (here placed in the subfamily Momphinae, revised status). High resolution photographs were taken to illustrate the habitus and various morphological structures (genitalia, notably) of the 24 above-mentioned type specimens (MNHN material).

Fourteen new species of the genus Thubdora Park, 2018 (Lepidoptera: Gelechioidea: Lecithoceridae) from Uganda, and three new combinations in Ptilothyris Walsingham, 1897 from DR Congo.

Fourteen new species of the Afrotropical genus Thubdora Park, 2018 (Lecithoceridae: Torodorinae) are described from Uganda: Thubdora afropyralidis Park, sp. nov., T. fruticosa Park, sp. nov., T. kibalensis Park, sp. nov., T. klenodes Park, sp. nov., T. latidiscalis Park, sp. nov., T. mirinae Park, sp. nov., T. nemorosa Park, sp. nov., T. retusivalva Park, sp. nov., T. umbratilis Park, sp. nov., and T. wooriana Park, sp. nov. in the aciphalla species-group (10 species); and T. corystos Park, sp. nov., T. narusia Park, sp. nov., T. trigonoides Park, sp. nov., and T. villosiphalla Park, sp. nov. in the acutalis species-group (four species). Four species are reported for the first time from Uganda: T. ambliodes Park, 2018, T. bilobella Park, 2018, T. ealaensis Park De Prins, 2019, and T. ghesquierei Park De Prins, 2019; and the females of T. ealaensis and T. ghesquierei are newly described, and their genitalia are illustrated. In addition, three species of Ptilothyris Walsingham, 1897 previously recorded from DR Congo are transferred to Thubdora: T. brachysema (Meyrick, 1938), comb. nov., T. crocophracta (Meyrick, 1938), comb. nov., and T. nemophorella (Ghesquière, 1940), comb. nov. Lectotypes for T. brachysema and T. crocophracta are designated herein. Identifications of the new species and corresponding females were confirmed based on the comparison of COI barcode sequences. Adults and genitalia of all the new species and the three species transferred from Ptilothyris are illustrated.